Cook pasta according to package directions (usually 1 minute less is perfect). Drain in a colander, rinse with cold water, and drain again; set aside.
In a large bowl, add tomatoes, cucumber, olives, onions, pepperoncinis, feta cheese, and fresh basil. Gently toss together and add the pasta.
For the dressing, place the red wine vinegar, olive oil, oregano, salt and pepper, fresh basil, lemon juice, garlic, and sugar in a jar with lid; shake well to mix.
Drizzle dressing over the salad; gently toss to coat. If you need to, add more olive oil. Cover and chill in refrigerator for 2-3 hours or overnight for the best flavor.
When ready to serve, toss the pasta salad again, garnish with fresh pepper and basil, and enjoy cold or at room temperature.